Extant: 21 valid subfamilies (21 unavailable), 66 valid tribes (62 synonyms), 308 valid genera (407 synonyms, 17 homonyms, 7 unavailable, 2 excluded from Formicidae), 12,748 valid species (2,223 synonyms, 213 homonyms, 4 unidentifiable, 110 unavailable, 3 excluded from Formicidae, 4,039 original combinations), 2,198 valid subspecies (2,162 synonyms, 104 homonyms, 783 unavailable, 890 original combinations)
Fossil: 4 valid subfamilies, 8 valid tribes (4 synonyms), 138 valid genera (16 synonyms, 5 homonyms, 1 unavailable, 5 excluded from Formicidae, 4 collective group names), 685 valid species (23 synonyms, 5 homonyms, 2 unavailable, 4 excluded from Formicidae, 3 collective group names), 5 valid subspecies
: 124 [Formicariae]; Leach, 1815Leach, W. E. 1815. Entomology. Pp. 57-172 in: Brewster, D. (ed.) 1815. The Edinburgh encyclopedia. Edinburgh: William Blackwood, Volume 9. [1815-04]
: 147 [Formicarides]; Stephens, 1829bStephens, J. F. 1829b. A systematic catalogue of British insects: being an attempt to arrange all the hitherto discovered indigenous insects in accordance with their natural affinities. London: Baldwin & Cradock, xxxiv + 416 + 388 pp. [1829-07-15]
: 356 [first spelling as Formicidae]; Haliday, 1836Haliday, A. H. 1836. Descriptions, etc. of the Hymenoptera. Pp. 316-331 in: Curtis, J.; Haliday, A. H.; Walker, F. 1836. Descriptions, etc. of the insects collected by Captain P. P. King R. N., F. R. S., in the survey of the Straits of Magellan. Transactions of the Linnean Society of London 17:315-359. [1836-06-21]
: 331; Westwood, 1839aWestwood, J. O. 1839a. An introduction to the modern classification of insects; founded on the natural habits and corresponding organisation of the different families. Volume 2. Part XI. London: Longman, Orme, Brown, Green and Longmans, pp. 193-224. [1839-03]
: 217; Swainson & Shuckard, 1840Swainson, W.; Shuckard, W. E. 1840. On the history and natural arrangement of insects. London: Longman, Brown, Green & Longman's, 406 pp. [1840-12] PDF
: 171; Nylander, 1846aNylander, W. 1846a. Adnotationes in monographiam formicarum borealium Europae. Acta Societatis Scientiarum Fennicae 2:875-944. [1846] PDF
: 877; Foerster, 1850aFoerster, A. 1850a. Hymenopterologische Studien. 1. Formicariae. Aachen: Ernst Ter Meer, 74 pp. [1850]
: 1 [Formicariae]; Mayr, 1855Mayr, G. 1855. Formicina austriaca. Beschreibung der bisher im österreichischen Kaiserstaate aufgefundenen Ameisen, nebst Hinzufügung jener in Deutschland, in der Schweiz und in Italien vorkommenden Arten. Verhandlungen der Zoologisch-Botanischen Vereins in Wien 5:273-478. [1855] PDF
: 275 [Formicina]; Smith, 1857aSmith, F. 1857a. Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A. R. Wallace. [part]. Journal and Proceedings of the Linnean Society of London. Zoology 2:42-88. [1857-11-02] PDF
: 52; Smith, 1858aSmith, F. 1858a. Catalogue of hymenopterous insects in the collection of the British Museum. Part VI. Formicidae. London: British Museum, 216 pp. [1858-03-27] PDF
: 1; Mayr, 1861Mayr, G. 1861. Die europäischen Formiciden. Nach der analytischen Methode bearbeitet. Wien: C. Gerolds Sohn, 80 pp. [1861] PDF
: 21; Mayr, 1865Mayr, G. 1865. Formicidae. In: Novara Expedition 1865. Reise der Österreichischen Fregatte "Novara" um die Erde in den Jahren 1857, 1858, 1859. Zoologischer Theil. Bd. II. Abt. 1. Wien: K. Gerold's Sohn, 119 pp. [1865-12-14]
: 6; Heer, 1867Heer, O. 1867. Fossile Hymenopteren aus Oeningen und Radoboj. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft für die Gesammten Naturwissenschaften 22(4):1-42. [1867] PDF
: 6 [Formicaria]; Forel, 1870Forel, A. 1870. Notices myrmécologiques. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 3:306-312. [1870-10] PDF
: 307 [Formicinae]; Forel, 1874Forel, A. 1874. Les fourmis de la Suisse. Systématique, notices anatomiques et physiologiques, architecture, distribution géographique, nouvelles expériences et observations de moeurs. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft für die Gesammten Naturwissenschaften 26:1-452. [1874] PDF
: 19 [Formicariae]; Dalla Torre, 1893: 1; Forel, 1899bForel, A. 1899b. Formicidae. Dummy reference. Biologia Centrali-Americana Hym 3:169.
: 1; Ruzsky, 1902dRuzsky, M. 1902d. Material on the ant fauna of the Caucasus and the Crimea. [In Russian.]. Protokoly Obshchestva Estestvoispytatelei pri Imperatorskom Kazanskom Universitete 206(suppl.):1-33. [1902]
: 5 [Formicarii]; Bingham, 1903Bingham, C. T. 1903. The fauna of British India, including Ceylon and Burma. Hymenoptera, Vol. II. Ants and Cuckoo-wasps. London: Taylor and Francis, 506 pp. [1903] PDF
: 1; Ruzsky, 1905bRuzsky, M. 1905b. The ants of Russia. (Formicariae Imperii Rossici). Systematics, geography and data on the biology of Russian ants. Part I. [In Russian.]. Trudy Obshchestva Estestvoispytatelei pri Imperatorskom Kazanskom Universitete 38(4-6):1-800. [1905]
: 91 [Formicariae or Formicidae]; Ashmead, 1905cAshmead, W. H. 1905c. A skeleton of a new arrangement of the families, subfamilies, tribes and genera of the ants, or the superfamily Formicoidea. Canadian Entomologist 37:381-384. [1905-11-01] PDF
: 384; all subsequent authors.
: 1 (Formicidae); Mayr, 1863aMayr, G. 1863a. Formicidarum index synonymicus. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 13:385-460. [1863] PDF
: 394 (Formicidae); Dalla Torre, 1893: 1 (Formicidae); Emery, 1910bEmery, C. 1910b. Hymenoptera. Fam. Formicidae. Subfam. Dorylinae. Genera Insectorum 102:1-34. [1910-06-25] PDF
: 3 (Dorylinae); Emery, 1911eEmery, C. 1911e. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118:1-125. [1911-09-09] PDF
: 2 (Ponerinae); Emery, 1913aEmery, C. 1913a ("1912"). Hymenoptera. Fam. Formicidae. Subfam. Dolichoderinae. Genera Insectorum 137:1-50. [1913-03-08] PDF
: 2 (Dolichoderinae); Emery, 1921cEmery, C. 1921c. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part]. Genera Insectorum 174A:1-94 + 7 plates. [1921-06-01]
: 3, Emery, 1922cEmery, C. 1922c. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part]. Genera Insectorum 174B:95-206. [1922-09-02] PDF
: 95 and Emery, 1924fEmery, C. 1924f ("1922"). Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [concl.]. Genera Insectorum 174C:207-397. [1924-06-24] PDF
: 207 (Myrmicinae); Emery, 1925dEmery, C. 1925d. Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum 183:1-302. [1925-12-31] PDF
: 2 (Formicinae); Shattuck, 1994Shattuck, S. O. 1994. Taxonomic catalog of the ant subfamilies Aneuretinae and Dolichoderinae (Hymenoptera: Formicidae). University of California Publications in Entomology 112:i-xix, 1-241. [1994-01] PDF
: 1 (Aneuretinae and Dolichoderinae); Bolton, 1995bBolton, B. 1995b. A new general catalogue of the ants of the world. Cambridge, Mass.: Harvard University Press, 504 pp. [1995-10]
: 7 (Formicidae)
: 161 (Formicidae); Emery, 1895lEmery, C. 1895l. Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 8:685-778. [1895-10-18] PDF
: 764 (Formicidae); Emery, 1896eEmery, C. 1896e. Clef analytique des genres de la famille des Formicides, pour la détermination des neutres. Annales de la Société Entomologique de Belgique 40:172-189. [1896-05-02>] PDF
: 173 (Formicidae); Ashmead, 1905cAshmead, W. H. 1905c. A skeleton of a new arrangement of the families, subfamilies, tribes and genera of the ants, or the superfamily Formicoidea. Canadian Entomologist 37:381-384. [1905-11-01] PDF
: 381 (Formicidae); Wheeler, 1910aWheeler, W. M. 1910a. Ants: their structure, development and behavior. New York: Columbia University Press, xxv + 663 pp. [1910-03] PDF
: 134 (Formicidae); Emery, 1910bEmery, C. 1910b. Hymenoptera. Fam. Formicidae. Subfam. Dorylinae. Genera Insectorum 102:1-34. [1910-06-25] PDF
: 3 (Dorylinae); Emery, 1911eEmery, C. 1911e. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118:1-125. [1911-09-09] PDF
: 2 (Ponerinae); Emery, 1913aEmery, C. 1913a ("1912"). Hymenoptera. Fam. Formicidae. Subfam. Dolichoderinae. Genera Insectorum 137:1-50. [1913-03-08] PDF
: 2 (Dolichoderinae); Forel, 1917Forel, A. 1917. Cadre synoptique actuel de la faune universelle des fourmis. Bulletin de la Société Vaudoise des Sciences Naturelles 51:229-253. [1917-03-08] PDF
: 235 (Formicidae); Emery, 1921cEmery, C. 1921c. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part]. Genera Insectorum 174A:1-94 + 7 plates. [1921-06-01]
: 3 and Emery, 1922cEmery, C. 1922c. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part]. Genera Insectorum 174B:95-206. [1922-09-02] PDF
: 95 (Myrmicinae); Wheeler, 1922Wheeler, W. M. 1922. Ants of the American Museum Congo expedition. New York: Bulletin of the American Museum of Natural History, 1139.
: 631 (Formicidae); Emery, 1924fEmery, C. 1924f ("1922"). Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [concl.]. Genera Insectorum 174C:207-397. [1924-06-24] PDF
: 207 (Myrmicinae); Emery, 1925dEmery, C. 1925d. Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum 183:1-302. [1925-12-31] PDF
: 2 (Formicinae); Brown, 1973b: 165, 178 (Formicidae); Snelling, 1981Snelling, R. R. 1981. Systematics of social Hymenoptera. Pp. 369-453 in: Hermann, H. R. (ed.) 1981. Social insects. Volume 2. New York: Academic Press, xiii + 491 pp. [1981] [1981]
: 387 (Formicidae); Wheeler & Wheeler, 1985bWheeler, G. C.; Wheeler, J. 1985b. A simplified conspectus of the Formicidae. Transactions of the American Entomological Society 111:255-264. [1985-07-30] PDF
: 256 (Formicidae); Dlussky & Fedoseeva, 1988Dlussky, G. M.; Fedoseeva, E. B. 1988. Origin and early stages of evolution in ants. [In Russian.]. Pp. 70-144 in: Ponomarenko, A. G. (ed.) 1988. Cretaceous biocenotic crisis and insect evolution. [In Russian.]. Moskva: Nauka, 232 pp. [1987-12-22>]
: 77 (Formicidae); Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 9 (Formicidae); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 12 (Formicidae); Bolton, 2003Bolton, B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71:1-370. [2003-11-10] PDF
: 4 (Formicidae); Engel & Grimaldi, 2005Engel, M. S.; Grimaldi, D. A. 2005. Primitive new ants in Cretaceous amber from Myanmar, New Jersey, and Canada (Hymenoptera: Formicidae). American Museum Novitates 3485:1-23. [2005-07-25] PDF
: 4 (Formicidae); Ward, 2007CWard, P. S. 2007C. Phylogeny, classification, and species-level taxonomy of ants (Hymenoptera: Formicidae). Zootaxa 1668:549-563. [2007-12-21] PDF
: 549 (Formicidae)
: 173 (Formicidae subfamilies and genera); Emery, 1910bEmery, C. 1910b. Hymenoptera. Fam. Formicidae. Subfam. Dorylinae. Genera Insectorum 102:1-34. [1910-06-25] PDF
: 4 (Dorylinae tribes and genera); Emery, 1911eEmery, C. 1911e. Hymenoptera. Fam. Formicidae. Subfam. Ponerinae. Genera Insectorum 118:1-125. [1911-09-09] PDF
: 4 (Ponerinae tribes and genera); Emery, 1913aEmery, C. 1913a ("1912"). Hymenoptera. Fam. Formicidae. Subfam. Dolichoderinae. Genera Insectorum 137:1-50. [1913-03-08] PDF
: 6 (Dolichoderinae tribes and genera); Escherich, 1917Escherich, K. 1917. Die Ameise, Schilderung ihrer Lebensweise. Zweite verbesserte und vermehrte Auflage. Braunschweig: Friedrich Vieweg und Sohn, xvi + 348 pp. [1917]
: 2 (Formicidae subfamilies); Emery, 1921cEmery, C. 1921c. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part]. Genera Insectorum 174A:1-94 + 7 plates. [1921-06-01]
: 9, 12, 16 (Myrmicinae tribes and genera); Wheeler, 1922Wheeler, W. M. 1922. Ants of the American Museum Congo expedition. New York: Bulletin of the American Museum of Natural History, 1139.
: 631 (Formicidae subfamilies, tribes and genera); Emery, 1925dEmery, C. 1925d. Hymenoptera. Fam. Formicidae. Subfam. Formicinae. Genera Insectorum 183:1-302. [1925-12-31] PDF
: 7 (Formicinae tribes and genera); Clark, 1951Clark, J. 1951. The Formicidae of Australia. Vol. 1. Subfamily Myrmeciinae. Melbourne: CSIRO, 230 pp. [1951] PDF
: 14 (Formicidae subfamilies); Wheeler & Wheeler, 1972aWheeler, G. C.; Wheeler, J. 1972a. The subfamilies of Formicidae. Proceedings of the Entomological Society of Washington 74:35-45. [1972-03-28] PDF
: 42 (Formicidae subfamilies); Snelling, 1981Snelling, R. R. 1981. Systematics of social Hymenoptera. Pp. 369-453 in: Hermann, H. R. (ed.) 1981. Social insects. Volume 2. New York: Academic Press, xiii + 491 pp. [1981] [1981]
: 386 (Formicidae subfamilies); Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 33 (Formicidae subfamilies and genera); Bolton, 1990cBolton, B. 1990c. Army ants reassessed: the phylogeny and classification of the doryline section (Hymenoptera, Formicidae). Journal of Natural History 24:1339-1364. [1990-10-19] PDF
: 1361 (Formicidae subfamilies); Shattuck, 1992cShattuck, S. O. 1992c. Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology 21:1-181. [1992] PDF
: 20 (Dolichoderinae genera); Ward, 1990Ward, P. S. 1990. The ant subfamily Pseudomyrmecinae (Hymenoptera: Formicidae): generic revision and relationship to other formicids. Systematic Entomology 15:449-489. [1990-11] PDF
: 464 (Pseudomyrmecinae genera); Brothers & Finnamore, 1993Brothers, D. J.; Finnamore, A. T. 1993. Superfamily Vespoidea. Pp. 161-278 in: Goulet, H.; Huber, J. T. (eds.) 1993. Hymenoptera of the world: an identification guide to families. Research Branch, Agriculture Canada Publication 1894/E. Ottawa: Canada Communications Group, vii + 668 pp. [1993]
: 218 (Formicidae subfamilies); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 7 (Formicidae subfamilies and genera); Bolton, 2003Bolton, B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71:1-370. [2003-11-10] PDF
: 11 (revision of subfamilies)
: 339 (proventriculus); Janet, 1902: 25 (general anatomy); Pietschker, 1910: 1 (brain structure); Wheeler, 1910aWheeler, W. M. 1910a. Ants: their structure, development and behavior. New York: Columbia University Press, xxv + 663 pp. [1910-03] PDF
: 13 (general morphology, anatomy); Emery, 1913dEmery, C. 1913d. La nervulation de l'aile antérieure des Formicides. Revue Suisse de Zoologie 21:577-587. [1913-09] PDF
: 577 (venation); Wheeler, 1918cWheeler, W. M. 1918c. A study of some ant larvae, with a consideration of the origin and meaning of the social habit among insects. Proceedings of the American Philosophical Society 57:293-343. [1918-08-08] PDF
: 293 (larvae); Janet, 1923: 5 (anatomical drawings synopsis); Bugnion, 1930Bugnion, E. 1930. Les pièces buccales, le sac infrabuccal et le pharynx des fourmis. Bulletin. Société Entomologique d'Egypte 14:85-210. [1930]
: 85 (mouthparts); Brown & Nutting, 1950: 113 (venation); Gregg, 1953cGregg, R. E. 1953c. Morphological considerations affecting the taxonomy of certain genera of ants (Hymenoptera, Formicidae). Proceedings of the Entomological Society of Washington 55:324-330. [1953-12-29] PDF
: 328 (promesonotal suture motility); Eisner, 1957Eisner, T. 1957. A comparative morphological study of the proventriculus of ants (Hymenoptera: Formicidae). Bulletin of the Museum of Comparative Zoology 116:439-490. [1957-07] PDF
: 439 (proventriculus); Gotwald, 1969: 1 (mouthparts and gaster); Crozier, 1970aCrozier, R. H. 1970a. Karyotypes of twenty-one ant species (Hymenoptera: Formicidae), with reviews of the known ant karyotypes. Canadian Journal of Genetics and Cytology 12:109-128. [1970-03] PDF
: 109 (karyology); Markl, 1973Markl, H. 1973. The evolution of stridulatory communication in ants. Pp. 258-265 in: International Union for the Study of Social Insects. Congress 1973. Proceedings IUSSI VIIth International Congress, London, 10-15 September, 1973. Southampton: University of Southampton, vi + 418 pp. [1973]
: 258 (stridulitrum); Wheeler & Wheeler, 1976bWheeler, G. C.; Wheeler, J. 1976b. Ant larvae: review and synthesis. Memoirs of the Entomological Society of Washington 7:1-108. [1976] PDF
: 45 (larvae); Hölldobler & Engel, 1979: 285 and Hölldobler & Engel-Siegel, 1982Hölldobler, B.; Engel-Siegel, H. 1982. Tergal and sternal glands in male ants. Psyche (Cambridge) 89:113-132. [1982-12-17]
: 113 (tergal and sternal glands); Imai, Baroni Urbani, Kubota, et al. 1984: 5 (karyology); Imai, Brown, Kubota, et al. 1984: 66 (karyology); Imai, Kubota, Brown, et al. 1985: 46 (karyology); Hölldobler & Engel-Siegel, 1985Hölldobler, B.; Engel-Siegel, H. 1985 ("1984"). On the metapleural gland of ants. Psyche (Cambridge) 91:201-224. [1985-04-22] PDF
: 201 (metapleural gland); Billen, 1986bBillen, J. P. J. 1986b. Comparative morphology and ultrastructure of the Dufour gland in ants (Hymenoptera: Formicidae). Entomologia Generalis 11:165-181. [1986-05-30]
: 165 (Dufour's gland); Wheeler & Wheeler, 1986eWheeler, G. C.; Wheeler, J. 1986e. Ten-year supplement to "Ant larvae: review and synthesis". Proceedings of the Entomological Society of Washington 88:684-702. [1986-10-14] PDF
: 684 (larvae); Brown, 1988h: 17 (malpighian tubule numbers); Caetano, 1988Caetano, F. H. 1988. Anatomia, histologia e histoquímica do sistema digestivo e excretor de operárias de formigas (Hymenoptera, Formicidae). Naturalia (São Paulo) 13:129-174. [1988]
: 129 (digestive tract anatomy); Francoeur & Loiselle, 1988aFrancoeur, A.; Loiselle, R. 1988a. Évolution du strigile chez les Formicides (Hyménoptères). Naturaliste Canadien (Québec) 115:333-353. [1988-12] PDF
: 333 (strigil, evolution); Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 5, 229 (anatomy, exocrine glands); Hashimoto, 1991aHashimoto, Y. 1991a. Phylogenetic study of the family Formicidae based on the sensillum structures on the antennae and labial palpi (Hymenoptera, Aculeata). Japanese Journal of Entomology 59:125-140. [1991-03-25] PDF
: 125 (antennal and labial sensilla); Hashimoto, 1991bHashimoto, Y. 1991b. Phylogenetic implications of the spur structures of the hind tibia in the Formicidae (Hymenoptera). Japanese Journal of Entomology 59:289-294. [1991-06-25] PDF
: 289 (tibial spurs); Billen, 1993Billen, J. P. J. 1993. Morphology of the exocrine system in ants. Materialy Kollokviumov Sektsii Obshchestvennykh Nasekomykh Vsesoyuznogo Entomologicheskogo Obshchestva 2:1-15.
: 4 (exocrine glands); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 191 (morphology glossary); Hölldobler, Obermayer & Peeters, 1996: 157 (metatibial gland); Wenseleers, Schoeters, et al. 1998: 121 (cloacal gland); Perrault, 1999APerrault, G. H. 1999A ("1998"). Heteroponera georgesi n. sp. de Guyane française (Hymenoptera, Formicidae). Bulletin de la Société Entomologique de France 103:475-477. [1999-01-22] PDF
: 125 (thoracic anatomy); Perrault, 2005Perrault, G. H. 2005 ("2004"). Étude morphoanatomique et biométrique du métasoma antérieur des ouvrières. Contribution à la systématique et à la phylogénie des fourmis (Hymenoptera: Formicidae). Annales de la Société Entomologique de France (n.s.)40:291-371. [2005-08-10]
: 291 (anterior metasomal anatomy); Serna & MacKay, 2010Serna, F.; MacKay, W. 2010. A descriptive morphology of the ant genus Procryptocerus (Hymenoptera: Formicidae). Journal of Insect Science 10: Article 111 (available online: insectscience.org/10.111). PDF
: 1 (Procryptocerus morphology); Keller, 2011Keller, R. A. 2011. A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the poneromorph subfamilies. Bulletin of the American Museum of Natural History 355:1-90. [2011-08-10] PDF
: 1 (phylogenetic analysis of morphology)
: 52; Emery, 1920eEmery, C. 1920e. La distribuzione geografica attuale delle formiche. Tentativo di spiegarne la genesi col soccorso di ipotesi filogenetiche e paleogeografiche. Atti della Reale Accademia dei Lincei. Memorie. Classe di Scienze Fisiche, Matematiche e Naturali (5)13:357-450. [1920]
: 370; Donisthorpe, 1922bDonisthorpe, H. 1922b ("1921"). The subfamilies of Formicidae. Transactions of the Entomological Society of London 1921:xl-xlvi. [1922-04-13] PDF
: xlvi; Wheeler, 1923gWheeler, W. M. 1923g. Social life among the insects. New York: Harcourt, Brace and Co., vii + 375 pp. [1923]
: 156; Wheeler, 1928hWheeler, W. M. 1928h. The social insects: their origin and evolution. New York: Harcourt, Brace and Co., xviii + 378 pp. [1928]
: 124; Stitz, 1935Stitz, H. 1935. Formicidae. Dummy reference. Tabulae Biologicae Periodicae 5:208-280.
: 211; Morley, 1938Morley, B. D. W. 1938. An outline of the phylogeny of the Formicidae. Bulletin de la Société Entomologique de France 43:190-194. [1938-10-05] PDF
: 190; Brown & Nutting, 1950: 113; Bernard, 1951cBernard, F. 1951c. Hyménoptères, super-famille des Formicoidea Ashmead 1905. Pp. 997-1104 in: Grassé, P. P. (ed.) 1951. Traité de Zoologie. Tome X. Fasc. II. Paris: Masson et Cie, pp. 976-1948. [1951]
: 1041; Brown, 1954e: 30; Wilson, Carpenter & Brown, 1967: 12; Taylor, 1978aTaylor, R. W. 1978a. Nothomyrmecia macrops: a living-fossil ant rediscovered. Science (Washington, D. C.) 201:979-985. [1978-09-15] PDF
: 982; Rasnitsyn, 1980Rasnitsyn, A. P. 1980. Origin and evolution of Hymenoptera. [In Russian.]. Trudy Paleontologicheskogo Instituta. Akademiya Nauk SSSR 174:1-192. [1980-01-24>]
: 45-46 (foldout page); Lutz, 1986Lutz, H. 1986. Eine neue Unterfamilie der Formicidae (Insecta: Hymenoptera) aus dem mittel-eozänen Olschiefer der "Grube Messel" bei Darmstadt (Deutschland, S-Hessen). Senckenbergiana Lethaea 67:177-218. [1986-10-31] PDF
: 213; Dlussky & Fedoseeva, 1988Dlussky, G. M.; Fedoseeva, E. B. 1988. Origin and early stages of evolution in ants. [In Russian.]. Pp. 70-144 in: Ponomarenko, A. G. (ed.) 1988. Cretaceous biocenotic crisis and insect evolution. [In Russian.]. Moskva: Nauka, 232 pp. [1987-12-22>]
: 137; Baroni Urbani, 1989Baroni Urbani, C. 1989. Phylogeny and behavioural evolution in ants, with a discussion of the role of behaviour in evolutionary processes. Ethology Ecology and Evolution 1:137-168. [1989-10] PDF
: 137; Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 26; Hashimoto, 1991aHashimoto, Y. 1991a. Phylogenetic study of the family Formicidae based on the sensillum structures on the antennae and labial palpi (Hymenoptera, Aculeata). Japanese Journal of Entomology 59:125-140. [1991-03-25] PDF
: 125; Hashimoto, 1991bHashimoto, Y. 1991b. Phylogenetic implications of the spur structures of the hind tibia in the Formicidae (Hymenoptera). Japanese Journal of Entomology 59:289-294. [1991-06-25] PDF
: 289; Baroni Urbani, Bolton & Ward, 1992Baroni Urbani, C.; Bolton, B.; Ward, P. S. 1992. The internal phylogeny of ants (Hymenoptera: Formicidae). Systematic Entomology 17:301-329. [1992-10] PDF
: 301; Shattuck, 1992bShattuck, S. O. 1992b. Higher classification of the ant subfamilies Aneuretinae, Dolichoderinae and Formicinae (Hymenoptera: Formicidae). Systematic Entomology 17:199-206. [1992-04] PDF
: 203; Brothers & Carpenter, 1993Brothers, D. J.; Carpenter, J. M. 1993. Phylogeny of Aculeata: Chrysidoidea and Vespoidea (Hymenoptera). Journal of Hymenoptera Research 2:227-304. [1993-09-30] PDF
: 227; Ward, 1994Ward, P. S. 1994. Adetomyrma, an enigmatic new ant genus from Madagascar (Hymenoptera: Formicidae), and its implications for ant phylogeny. Systematic Entomology 19:159-175. [1994-08-01] PDF
: 173; Hashimoto, 1996Hashimoto, Y. 1996. Skeletomuscular modifications associated with the formation of an additional petiole on the anterior abdominal segments in aculeate Hymenoptera. Japanese Journal of Entomology 64:340-356. [1996-06-25] PDF
: 340; Crozier, Jermiin & Chiotis, 1997Crozier, R. H.; Jermiin, L. S.; Chiotis, M. 1997. Molecular evidence for a Jurassic origin of ants. Naturwissenschaften 84:22-23. [1997-01] PDF
: 22; Grimaldi, Agosti & Carpenter, 1997Grimaldi, D.; Agosti, D.; Carpenter, J. M. 1997. New and rediscovered primitive ants (Hymenoptera: Formicidae) in Cretaceous amber from New Jersey, and their phylogenetic relationships. American Museum Novitates 3208:1-43. [1977-10-23] PDF
: 25; Perrault, 1999APerrault, G. H. 1999A ("1998"). Heteroponera georgesi n. sp. de Guyane française (Hymenoptera, Formicidae). Bulletin de la Société Entomologique de France 103:475-477. [1999-01-22] PDF
: 125; Brothers, 1999Brothers, D. J. 1999. Phylogeny and evolution of wasps, ants and bees (Hymenoptera, Chrysidoidea, Vespoidea and Apoidea). Zoologica Scripta 28:233-249.
: 233; Grimaldi & Agosti, 2000BGrimaldi, D.; Agosti, D. 2000B. A formicine in New Jersey Cretaceous amber (Hymenoptera: Formicidae) and early evolution of the ants. Proceedings of the National Academy of Sciences of the United States of America 97:13678-13683. [2000-12-05]
: 13680; Ohnishi, Imai, et al. 2003: 419; Astruc, Julien, et al. 2004: 880; Grimaldi & Engel, 2005Grimaldi, D.; Engel, M. S. 2005. Evolution of the insects. Cambridge: Cambridge University Press, xv + 755 pp. PDF
: 440; Wilson & Hölldobler, 2005A+C1428Wilson, E. O.; Hölldobler, B. 2005A+C1428. The rise of the ants: a phylogenetic and ecological explanation. Proceedings of the National Academy of Sciences of the United States of America 102:7411-7414. [2005-05-24] PDF
: 7411; Brady, Schultz, et al. 2006: 18172; Moreau, Bell, et al. 2006: 101; Ouellette, Fisher, et al. 2006: 359; Rabeling, Brown & Verhaagh, 2008Rabeling, C.; Brown, J. M.; Verhaagh, M. 2008. Newly discovered sister lineage sheds light on early ant evolution. Proceedings of the National Academy of Sciences of the United States of America 105:14913-14917. [2008-09-30] PDF
: 14914
: 682 (fossil taxa catalogue); Handlirsch, 1907Handlirsch, A. 1907. Die fossilen Insekten und die Phylogenie der rezenten Formen. Ein Handbuch für Paläontologen und Zoologen. Lieferung 6. Leipzig: Wilhelm Engelmann, pp. 801-960. [1907-06]
: 859 (fossil taxa catalogue); Wheeler, 1915iWheeler, W. M. 1915i ("1914"). The ants of the Baltic Amber. Schriften der Physikalisch-Ökonomischen Gesellschaft zu Königsberg 55:1-142. [1915] PDF
: 1 (Baltic amber fauna synopsis); Burnham, 1979Burnham, L. 1979 ("1978"). Survey of social insects in the fossil record. Psyche (Cambridge) 85:85-133. [1979-01-26] PDF
: 108 (fossils in amber checklist); Keilbach, 1982Keilbach, R. 1982. Bibliographie und Liste der Arten tierischer Einschlüsse in fossilen Harzen sowie ihrer Aufbewahrungsorte. Teil 1. Deutsche Entomologische Zeitschrift (N.F.)29:129-286. [1982-03-15]
: 272 (fossils in amber checklist); Spahr, 1987Spahr, U. 1987. Ergänzungen und Berichtigungen zu R. Keilbach's Bibliographie und Liste der Bernsteinfossilien - Ordnung Hymenoptera. Stuttgarter Beiträge zur Naturkunde. Serie B (Geologie und Paläontologie) 127:1-121. [1987-03-16]
: 41 (fossils in amber checklist); Carpenter, 1992Carpenter, F. M. 1992. Treatise on invertebrate paleontology. Part R. Arthropoda 4. Volume 4. Superclass Hexapoda. Boulder: Geological Society of America, pp. i-ii, 279-655. [1992-12-09]
: 490 (fossil genera review); Bolton, 1995bBolton, B. 1995b. A new general catalogue of the ants of the world. Cambridge, Mass.: Harvard University Press, 504 pp. [1995-10]
: 4 (fossil taxa checklist, in catalogue); Grimaldi, Agosti & Carpenter, 1997Grimaldi, D.; Agosti, D.; Carpenter, J. M. 1997. New and rediscovered primitive ants (Hymenoptera: Formicidae) in Cretaceous amber from New Jersey, and their phylogenetic relationships. American Museum Novitates 3208:1-43. [1977-10-23] PDF
: 1 (Cretaceous genera in amber review); Dlussky, 1997ADlussky, G. M. 1997A. Genera of ants (Hymenoptera: Formicidae) from Baltic amber. [In Russian.]. Paleontologicheskii Zhurnal 1997(6):50-62. PDF
: 617 (Baltic amber genera key); Bolton, 2003Bolton, B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71:1-370. [2003-11-10] PDF
: 73 (fossil subfamilies synopsis); Dlussky & Rasnitsyn, 2007Dlussky, G. M.; Rasnitsyn, A. P. 2007. Paleontological record and stages of ant evolution. [In Russian.]. Uspekhi Sovremennoi Biologii 127:118-134. [2007] PDF
: 118 (Palaeontological record); Dlussky & Rasnitsyn, 2009Dlussky, G. M.; Rasnitsyn, A. P. 2009. Ants (Insecta: Vespida: Formicidae) in the Upper Eocene amber of central and eastern Europe. Paleontological Journal 43:1024-1042. [2009-12] PDF
: 1024 (European Upper Eocene amber, synopsis of species)
: 1038 (census of extant taxa); Ward, Bolton, Shattuck & Brown, 1996: 55 (bibliography of systematics); Brown, 2000: 46 (overview of genera); Brandão, 2000Brandão, C. R. F. 2000. Major regional and type collections of ants (Formicidae) of the world and sources for the identification of ant species. Pp. 172-185 in: Agosti, D.; Majer, J. D.; Alonso, L. E.; Schultz, T. R. (eds.) 2000. Ants. Standard methods for measuring and monitoring biodiversity. Washington: Smithsonian Institution Press, xix + 280 pp. [2000-11] [2000-11]
: 172 (list of major ant collections); Bolton, 2003Bolton, B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71:1-370. [2003-11-10] PDF
: 15, 77 (diagnosis, synopsis); Bolton, 2007ABolton, B. 2007A. How to conduct large-scale taxonomic revisions in Formicidae. Memoirs of the American Entomological Institute 80:52-71. [2007-09]
: 51 (method for taxonomic revisions); Taylor, 2007Taylor, R. W. 2007. Bloody funny wasps! Speculations on the evolution of eusociality in ants. Memoirs of the American Entomological Institute 80:580-609. [2007-09]
: 580 (eusociality in ants)
: 778; Smith, 1958cSmith, M. R. 1958c. Family Formicidae. Pp. 108-162 in: Krombein, K. V. (ed.) 1958. Hymenoptera of America north of Mexico. Synoptic catalogue. First supplement. United States Department of Agriculture. Agriculture Monograph 2(suppl. 1):1-305. [1958-10-08]
: 108 (first supplement to previous); Smith, 1967aSmith, M. R. 1967a. Family Formicidae. Pp. 343-374 in: Krombein, K. V.; Burks, B. D. (eds.) 1967. Hymenoptera of America north of Mexico. Synoptic catalog. Second supplement. United States Department of Agriculture. Agriculture Monograph 2(suppl. 2):1-584. [1967-02]
: 343 (second supplement); Smith, 1979Smith, D. R. 1979. Superfamily Formicoidea. Pp. 1323-1467 in: Krombein, K. V.; Hurd, P. D.; Smith, D. R.; Burks, B. D. (eds.) 1979. Catalog of Hymenoptera in America north of Mexico. Volume 2. Apocrita (Aculeata). Washington, D.C.: Smithsonian Institution Press, pp. i-xvi, 1199-2209. [1979]
: 1323
: 3; Brandão, 1991Brandão, C. R. F. 1991. Adendos ao catálogo abreviado das formigas da região Neotropical (Hymenoptera: Formicidae). Revista Brasileira de Entomologia 35:319-412. [1991-08-31] PDF
: 319 (supplement to previous)
: 727, 1005
: 9
: 5; Taylor, 1987aTaylor, R. W. 1987a. A checklist of the ants of Australia, New Caledonia and New Zealand (Hymenoptera: Formicidae). CSIRO (Commonwealth Scientific and Industrial Research Organization) Division of Entomology Report 41:1-92. [1987-06] PDF
: 1; Taylor, 1987bTaylor, R. W. 1987b. A checklist of the ants of Australia, New Caledonia and New Zealand (Hymenoptera: Formicidae). First supplement, 10 July, 1987. CSIRO (Commonwealth Scientific and Industrial Research Organization) Division of Entomology Report 41(Suppl.1):1-5. [1987-07]
: 1 (supplement to previous)
: 299 (Austria); Mayr, 1861Mayr, G. 1861. Die europäischen Formiciden. Nach der analytischen Methode bearbeitet. Wien: C. Gerolds Sohn, 80 pp. [1861] PDF
: 25 (Europe); André, 1874André, Ern. 1874. Description des fourmis d'Europe pour servir à l'étude des insectes myrmécophiles. Dummy reference. Revue et Magasin de Zoologie Pure et Appliquée 3:152-235.
: 167 (Europe); Forel, 1874Forel, A. 1874. Les fourmis de la Suisse. Systématique, notices anatomiques et physiologiques, architecture, distribution géographique, nouvelles expériences et observations de moeurs. Neue Denkschriften der Allgemeinen Schweizerischen Gesellschaft für die Gesammten Naturwissenschaften 26:1-452. [1874] PDF
: 19 (Switzerland); Saunders, 1880Saunders, E. 1880. Synopsis of the British Heterogyna and fossorial Hymenoptera. Transactions of the Entomological Society of London 1880:201-304. [1880-12-31] PDF
: 202 (Britain); André, 1882a: 125 (Europe & Algeria); Nasonov, 1889Nasonov, N. V. 1889. Contribution to the natural history of the ants primarily of Russia. 1. Contribution to the ant fauna of Russia. [In Russian.]. Izvestiya Imperatorskago Obshchestva Lyubitelei Estestvoznaniya Antropologii i Etnografii pri Imperatorskom Moskovskom Universitete 58:1-78. [1889]
: 50 (Russia); Lameere, 1892Lameere, A. 1892. Note sur les fourmis de la Belgique. Annales de la Société Entomologique de Belgique 36:61-69. [1892-02-06>]
: 62 (Belgium); Saunders, 1896Saunders, E. 1896. The Hymenoptera Aculeata of the British islands. A descriptive account of the families, genera, and species indigenous to Great Britain and Ireland, with notes as to habits, localities, habitats, etc. London: L. Reeve & Co., 391 pp. [1896] PDF
: 18 (Britain); Ruzsky, 1905bRuzsky, M. 1905b. The ants of Russia. (Formicariae Imperii Rossici). Systematics, geography and data on the biology of Russian ants. Part I. [In Russian.]. Trudy Obshchestva Estestvoispytatelei pri Imperatorskom Kazanskom Universitete 38(4-6):1-800. [1905]
: 99 (Russian Empire); Wasmann, 1906Wasmann, E. 1906. Zur Kenntniss der Ameisen und Ameisengäste von Luxemburg. Archives. Institut Grand-Ducal de Luxembourg (n.s.)1(1/2):104-124. [1906-06] PDF
: 7 (Luxemburg); Emery, 1908a (Palaearctic); Bondroit, 1910Bondroit, J. 1910 ("1909"). Les fourmis de Belgique. Annales de la Société Entomologique de Belgique 53:479-500. [1910-01-05] PDF
: 480 (Belgium); Stitz, 1914Stitz, H. 1914. Die Ameisen (Formicidae) Mitteleuropas, insbesondere Deutschlands. Pp. 1-111 in: Schröder, C. (ed.) 1914. Die Insekten Mitteleuropas insbesondere Deutschlands. Band II, Hymenopteren, 2. Teil. Stuttgart: Franckh'sche Verlagshandlung, 256 pp. [1914] PDF
: 54 (Central Europe); Donisthorpe, 1915fDonisthorpe, H. 1915f. British ants, their life-history and classification. Plymouth: Brendon & Son Ltd., xv + 379 pp. [1915]
: 65 (Britain); Forel, 1915dForel, A. 1915d. Fauna insectorum helvetiae. Hymenoptera. Formicidae. Die Ameisen der Schweiz. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 12(Beilage):1-77. [1915]
: 1 (Switzerland); Emery, 1916aEmery, C. 1916a ("1915"). Fauna entomologica italiana. I. Hymenoptera.-Formicidae. Bullettino della Società Entomologica Italiana 47:79-275. [1916-11-30] PDF
: 92 (Italy); Bondroit, 1918Bondroit, J. 1918. Les fourmis de France et de Belgique. Annales de la Société Entomologique de France 87:1-174. [1918-12-24] PDF
: 12 (France & Belgium); Kutter, 1920b: 132 (Switzerland); Soudek, 1922bSoudek, S. 1922b. Mravenci. Soustava, zemepisné rozsírení, oekologie a urcovací klíc mravencu zijících na území Ceskoslovenské republiky. Praha: Ceskoslovenská Spolecnost Entomologická, 100 pp. [1922] PDF
: 17 (Czechoslovakia); Lomnicki, 1925aLomnicki, J. 1925a ("1924"). Przeglad polskich gatunków rodzaju mrówki (Formica Linné). Polskie Pismo Entomologiczne 3:151-182. [1925-03-15<] PDF
: 160 (Poland); Stärcke, 1926: 79 (Netherlands); Karavaiev, 1927dKaravaiev, V. 1927d. The ant fauna of Ukraine. [In Ukrainian.]. Zbirnyk Prats' Zoolohichnoho Muzeyu 2:1-52 [= Trudy. Ukrains'ka Akademiya Nauk. Fizichno-Matematichnoho Viddilu 4:247-296]. [1927]
: 254 (Ukraine); Donisthorpe, 1927cDonisthorpe, H. 1927c. British ants, their life history and classification (2nd edn.). London: G. Routledge and Sons, xvi + 436 pp. [1927]
: 66 (Britain); Arnol'di, 1933aArnol'di, K. V. 1933a. Formicidae - ants. [In Russian.]. Pp. 594-605 in: Filip'ev, I. N.; Ogloblin, D. A. (eds.) 1933. Keys to insects. [In Russian.]. Moskva: Ogiz, viii + 820 pp. [1933-12-03>]
: 596 (Russia); Menozzi, 1933bMenozzi, C. 1933b. Le formiche della Palestina. Memorie della Società Entomologica Italiana 12:49-113. [1933-06-15] PDF
: 87 (Israel); Karavaiev, 1934Karavaiev, V. 1934. The fauna of the family Formicidae (ants) of the Ukraine. [In Ukrainian.]. Trudy Instytutu Zoolohii ta Biolohii Vseukrains'koi Akademii Nauk. Seriya 1. Pratsi z Systematyky ta Faunistyky 1934:1-164. [1934-09-27>]
: 48 (Ukraine); Stitz, 1939Stitz, H. 1939. Die Tierwelt Deutschlands und der angrenzenden Meersteile nach ihren Merkmalen und nach ihrer Lebensweise. 37. Theil. Hautflüger oder Hymenoptera. I: Ameisen oder Formicidae. Jena: G. Fischer, 428 pp. [1939]
: 56 (Germany); Kratochvíl, 1941bKratochvíl, J. 1941b. [Untitled. New subgenera of Tetramorium, and a new subspecies, Tetramorium ferox silhavyi, attributed to Kratochvíl.]. P. 84 in: Novák, V.; Sadil, J. 1941. Klíc k urcování mravencu strední Evropy se zvlástním zretelem k mravencí zvírene Cech a Moravy. Entomologické Listy 4:65-115. [1941]
: 69 (Central Europe); Novák & Sadil, 1941Novák, V.; Sadil, J. 1941. Klíc k urcování mravencu strední Evropy se zvlástním zretelem k mravencí zvírene Cech a Moravy. Entomologické Listy 4:65-115. PDF
: 69 (Central Europe); Ceballos, 1943Ceballos, G. 1943. Las tribus de los himenópteros de España. [part]. Madrid: Instituto Español de Entomología, pp. 289-420. [1943-12-30]
: 313 (Spain); Holgersen, 1943cHolgersen, H. 1943c. Bestemmelsestabell over norske maur (Hym., Formicidae). Norsk Entomologisk Tidsskrift 6:164-182. [1943-09-01] PDF
: 166 and Holgersen, 1944aHolgersen, H. 1944a. The ants of Norway (Hymenoptera, Formicidae). Nytt Magasin for Naturvidenskapene 84:165-203. [1944-09-08] PDF
: 198 (Norway); Boven, 1947: 168 (Belgium); Collingwood, 1958bCollingwood, C. A. 1958b. A key to the species of ants (Hymenoptera, Formicidae) found in Britain. Transactions of the Society for British Entomology 13:69-96. [1958-04-30] PDF
: 69 (Britain); Somfai, 1959Somfai, E. 1959. Hangya alkatúak Formicoidea. Magyarország Allatvilága 43:1-79. [1959-02-11]
: 1 (Hungary); Boven, 1959: 1 (Netherlands); Collingwood, 1964bCollingwood, C. A. 1964b. The identification and distribution of British ants. 1. A revised key to the species found in Britain. Transactions of the Society for British Entomology 16:93-114,121. [1964-12] PDF
: 93 (Britain); Bernard, 1967aBernard, F. 1967a ("1968"). Faune de l'Europe et du Bassin Méditerranéen. 3. Les fourmis (Hymenoptera Formicidae) d'Europe occidentale et septentrionale. Paris: Masson, 411 pp. [1967-11] PDF
: 75 (Western Europe); Boven, 1970b: 7 (Netherlands); Tarbinsky, 1976Tarbinsky, Y. S. 1976. The ants of Kirghizia. [In Russian.]. Frunze: Ilim, 217 pp. [1976-06-04>] PDF
: 14 (Kyrghyzstan); Bolton & Collingwood, 1975Bolton, B.; Collingwood, C. A. 1975. Handbooks for the identification of British Insects. Hymenoptera, Formicidae. Handbooks for the Identification of British Insects 6(3(c)):1-34. [1975-06]
: 1 (Britain); Boven, 1977: 1 (Belgium); Kutter, 1977cKutter, H. 1977c. Hymenoptera, Formicidae. Insecta Helvetica. Fauna 6:1-298. [1977]
: 19 (Switzerland); Brian, 1977Brian, M. V. 1977. Ants. London: Collins, 223 pp.
: 33 (Britain); Arnol'di & Dlussky, 1978Arnol'di, K. V.; Dlussky, G. M. 1978. Superfam. Formicoidea. 1. Fam. Formicidae - ants. [In Russian.]. Pp. 519-556 in: Medvedev, G. S. (ed.) 1978. Keys to the insects of the European part of the USSR. Vol. 3. Hymenoptera. Part 1. [In Russian.]. Opredeliteli Faune SSSR 119:3-584. [1978-11-02>]
: 520 (former European U.S.S.R.); Collingwood, 1978Collingwood, C. A. 1978. A provisional list of Iberian Formicidae with a key to the worker caste (Hym. Aculeata). EOS. Revista Española de Entomología 52:65-95. [1978-05-10] PDF
: 65 (Iberian Peninsula); Collingwood, 1979Collingwood, C. A. 1979. The Formicidae (Hymenoptera) of Fennoscandia and Denmark. Fauna Entomologica Scandinavica 8:1-174. [1979] PDF
: 28 (Fennoscandia & Denmark); Schembri & Collingwood, 1981Schembri, S. P.; Collingwood, C. A. 1981. A revision of the myrmecofauna of the Maltese Islands (Hymenoptera, Formicidae). Annali del Museo Civico di Storia Naturale "Giacomo Doria" 83:417-442. [1981-09-01] PDF
: 417 (Malta); Verhaeghe, Deligne, et al., 1984: 105 (Belgium genera); Gösswald, 1985Gösswald, K. 1985. Organisation und Leben der Ameisen. Stuttgart: Wissenschaftliche Verlagsgesellschaft mbH, 355 pp. [1985]
: 262 (Germany); Collingwood, 1985Collingwood, C. A. 1985. Hymenoptera: Fam. Formicidae of Saudi Arabia. Fauna of Saudi Arabia 7:230-302. [1985] PDF
: 230 (Saudi Arabia); Nilsson & Douwes, 1987Nilsson, O.; Douwes, P. 1987. Norrländska myror med bestämningstabell till arbetarna. Natur i Norr 6:49-90.
: 56 (Norway); Agosti & Collingwood, 1987bAgosti, D.; Collingwood, C. A. 1987b. A provisional list of the Balkan ants (Hym. Formicidae) with a key to the worker caste. II. Key to the worker caste, including the European species without the Iberian. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 60:261-293. [1987-12-15] PDF
: 261 (Balkans); Casevitz-Weulersse, 1990bCasevitz-Weulersse, J. 1990b. Étude systématique de la myrmécofaune corse (Hymenoptera, Formicidae) (Première partie). Bulletin du Muséum National d'Histoire Naturelle. Section A. Zoologie, Biologie et Écologie Animales (4)12:135-163. [1990-08-02]
: 135 and Casevitz-Weulersse, 1990cCasevitz-Weulersse, J. 1990c. Étude systématique de la myrmécofaune corse (Hymenoptera, Formicidae) (Deuxième partie). Bulletin du Muséum National d'Histoire Naturelle. Section A. Zoologie, Biologie et Écologie Animales (4)12:415-442. [1990-11-24] PDF
: 415 (Corsica); Dlussky, Soyunov & Zabelin, 1990Dlussky, G. M.; Soyunov, O. S.; Zabelin, S. I. 1990. Ants of Turkmenistan. [In Russian.]. Ashkabad: Ylym Press, 273 pp. PDF
: 99 (Turkmenistan); Kupyanskaya, 1990aKupyanskaya, A. N. 1990a. Ants of the Far Eastern USSR. [In Russian.]. Vladivostok: Akademiya Nauk SSSR, 258 pp. [1990-09-13>]
: 83 (Far Eastern Russia); Bolton, in Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 34 (Palaearctic genera key); Atanasov & Dlussky, 1992: 47 (Bulgaria); Arakelian, 1994Arakelian, G. R. 1994. Fauna of the Republic of Armenia. Hymenopterous insects. Ants (Formicidae). [In Russian.]. Erevan: Gitutium, 153 pp. [1994-06-20>] PDF
: 1 (Armenia); Radchenko, 1994bRadchenko, A. G. 1994b. Identification table for the ants (Hymenoptera, Formicidae) of southern Siberia. [In Russian.]. Pp. 95-118 in: Kotenko, A. G. (ed.) 1994. Hymenopterous insects of Siberia and the Far East. [In Russian.]. Trudy Zapovednika "Daurskii" 3:1-147. [1994-06-10>]
: 95 (South Siberia); Seifert, 1994aSeifert, B. 1994a ("1993"). Die freilebenden Ameisenarten Deutschlands (Hymenoptera: Formicidae) und Angaben zu deren Taxonomie und Verbreitung. Abhandlungen und Berichte des Naturkundemuseums Görlitz 67(3):1-44. [1994-01-10] PDF
: 1 (Germany, synopsis); Onoyama & Terayama, 1994Onoyama, K.; Terayama, M. 1994. A list of references on Japanese ants. [In Japanese.]. Tokyo: Myrmecological Society of Japan, iii + 50 pp. [1994-03-25]
: 1 (Japan bibliography); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 22, 42, 75, 153 (Dolichoderinae, Formicinae, Myrmicinae, Ponerinae genera keys); Douwes, 1995Douwes, P. 1995. Sveriges myror. Entomologisk Tidskrift 116:83-99. [1995-11]
: 83 (Sweden); Kupyanskaya, 1995: 327 (Far Eastern Russia); Wu & Wang, 1995aWu, J.; Wang, C. 1995a. The ants of China. [In Chinese.]. Beijing: China Forestry Publishing House, x + 214 pp. [1995-06]
: 30 (China); Fontenla Rizo, 1995bFontenla Rizo, J. L. 1995b. Reflexiones sobre las hormigas "vagabundas" de Cuba. Cocuyo (Havana) 3:11-22. [1995-06]
: 16 (Cuba, tramp species key); Collingwood & Agosti, 1996Collingwood, C. A.; Agosti, D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna of Saudi Arabia 15:300-385. [1996-04-15] PDF
: 300 (Saudi Arabia); Seifert, 1996BSeifert, B. 1996B. Ameisen beobachten, bestimmen. Augsburg: Naturbuch Verlag, 351 pp. [1996-12-31]
: 106 (Central Europe); Skinner & Allen, 1996Skinner, G. J.; Allen, G. W. 1996. Ants. Naturalists' Handbook 24. Slough: Richmond Publishing Co., ii + 83 pp. [1996]
: 40 (Britain); Dlussky, 1997ADlussky, G. M. 1997A. Genera of ants (Hymenoptera: Formicidae) from Baltic amber. [In Russian.]. Paleontologicheskii Zhurnal 1997(6):50-62. PDF
: 616 (*Baltic Amber genera); Collingwood & Prince, 1998Collingwood, C.; Prince, A. 1998. A guide to ants of continental Portugal (Hymenoptera: Formicidae). Boletim da Sociedade Portuguesa de Entomologia. Suplemento 5:1-49. [1998-07]
: 9 (Portugal); Czechowski, Radchenko & Czechowska, 2002Czechowski, W.; Radchenko, A.; Czechowska, W. 2002. The ants (Hymenoptera, Formicidae) of Poland. Warsaw: Museum and Institute of Zoology, Polish Academy of Sciences, 200 pp. PDF
: 133 (Poland); Aktaç & Radchenko, 2002Aktaç, N.; Radchenko, A. G. 2002. Türkiye karincalarinin (Hymenoptera: Formicidae) altfamilya ve cins tani anahtarlari. Türkiye Entomoloji Dergisi 26:51-61. [2002-03]
: 53 (Turkey genera); Yoshimura & Onoyama, 2002BYoshimura, M.; Onoyama, K. 2002B. Male-based keys to the subfamilies and genera of Japanese ants (Hymenoptera: Formicidae). Entomological Science 5:421-443. [2002-12-25]
: 424 (Japan genera, males key); Imai, Kihara, Kondoh, et al. 2003: 8 (Japan); Radchenko, 2005b: 183 (North Korea); Seifert, 2007Seifert, B. 2007. Die Ameisen Mittel- und Nordeuropas. Tauer: lutra Verlags- und Vertriebsgesellschaft, 368 pp. [2007-04]
: 107 (North and Central Europe); Boer, 2010Boer, P. 2010. Mieren van de Benelux. ′s-Graveland, Netherlands: Stichting Jeugdbondsuitgeverij, 183 pp.
: 8 (Benelux); Lapeva-Gjonova, et al. 2010: 1 (Bulgaria species catalogue); Csősz, Markó & Gallé, 2011Csősz, S.; Markó, B.; Gallé, L. 2011. The myrmecofauna (Hymenoptera: Formicidae) of Hungary: an updated checklist. North-Western Journal of Zoology 7:55-62. [2011-06] PDF
: pagination? (Hungary species catalogue); Czechowski, Radchenko, et al. 2012: 333 (Poland)
: 596 (Canada); Provancher, 1887Provancher, L. 1887. Fam. X. - Les Formicides. Dummy reference. Pp. 224-249 in: Provancher, L. 1887. 1885-1889. Additions et corrections au Volume II de la Faune entomologique du Canada, traitant des Hyménoptères. Québec: C. Darveau, 477 pp.
: 225 (Canada); Cresson, 1887Cresson, E. T. 1887. Synopsis of the families and genera of the Hymenoptera of America, north of Mexico, together with a catalogue of the described species, and bibliography. Transactions of the American Entomological Society, Suppl. Vol. 1887:1-351. [1887]
: 93 (U.S.A.); Wheeler, 1910aWheeler, W. M. 1910a. Ants: their structure, development and behavior. New York: Columbia University Press, xxv + 663 pp. [1910-03] PDF
: 557 (North America genera); Wheeler, 1916rWheeler, W. M. 1916r. Formicoidea. Formicidae. Pp. 577-601 in: Viereck, H. L. 1916. Guide to the insects of Connecticut. Part III. The Hymenoptera, or wasp-like insects, of Connecticut. Connecticut State Geological and Natural History Survey. Bulletin 22:1-824. [1916]
: 579 (U.S.A., Connecticut); Cole, 1942: 359 (U.S.A., Utah); Smith, 1943eSmith, M. R. 1943e. A generic and subgeneric synopsis of the male ants of the United States. American Midland Naturalist 30:273-321. [1943-10-22] PDF
: 276 (U.S.A. subfamilies & genera, males); Buren, 1944aBuren, W. F. 1944a. A list of Iowa ants. Iowa State College Journal of Science 18:277-312. [1944-04] PDF
: 278 (U.S.A., Iowa); Smith, 1947fSmith, M. R. 1947f. A generic and subgeneric synopsis of the United States ants, based on the workers. American Midland Naturalist 37:521-647. [1947-07-27] PDF
: 524 (U.S.A. genera); Creighton, 1950aCreighton, W. S. 1950a. The ants of North America. Bulletin of the Museum of Comparative Zoology 104:1-585. [1950-04] PDF
: 29 (North America); Gregg, 1963Gregg, R. E. 1963. The ants of Colorado, with reference to their ecology, taxonomy, and geographic distribution. Boulder: University of Colorado Press, xvi + 792 pp. [1963]
: 278 (U.S.A., Colorado); Wheeler & Wheeler, 1963Wheeler, G. C.; Wheeler, J. 1963. The ants of North Dakota. Grand Forks, North Dakota: University of North Dakota Press, viii + 326 pp. [1963]
: 149 (U.S.A., North Dakota); Francoeur, 1979aFrancoeur, A. 1979a. Les fourmis du Québec. 1. Introduction. 2. La famille des Formicidae. 3. La sous-famille des Ponerinae. Annales de la Société Entomologique du Québec 24:12-47. [1979-01] PDF
: 48 (Canada, Québec, introduction); Allred, 1982Allred, D. M. 1982. Ants of Utah. Great Basin Naturalist 42:415-511. [1982-12-31]
: 437 (U.S.A., Utah); Wheeler & Wheeler, 1986gWheeler, G. C.; Wheeler, J. 1986g. The ants of Nevada. Los Angeles: Natural History Museum of Los Angeles County, vii + 138 pp. [1986] PDF
: 16 (U.S.A., Nevada); Mackay, Lowrie, et al., 1988: 81 (U.S.A., New Mexico); Cover, in Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 62 (Nearctic genera key); Mackay & MacKay, 2002Mackay, W.; MacKay, E. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Lewiston, New York: Edwin Mellen Press, 400 pp. [2002-12] PDF
: 16 (U.S.A., New Mexico); Dlussky & Rasnitsyn, 2003Dlussky, G. M.; Rasnitsyn, A. P. 2003 ("2002"). Ants (Hymenoptera: Formicidae) of Formation Green River and some other Middle Eocene deposits of North America. Russian Entomological Journal 11:411-436. [2003-06] PDF
: 430 (*Eocene North American genera); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 25, 38, 48, 95, 160 (Dolichoderinae, Ecitoninae, Formicinae, Myrmicinae, Ponerinae genera keys); Coovert, 2005: 1 (U.S.A., Ohio); Ward, 2005Ward, P. S. 2005. A synoptic review of the ants of California (Hymenoptera: Formicidae). Zootaxa 936:1-68. [2005-04-12] PDF
: 20 (U.S.A., California genera); Fisher & Cover, 2007Fisher, B. L.; Cover, S. P. 2007. Ants of North America. A guide to genera. Berkeley: University of California Press, xiv + 194 pp. [2007-09]
: 1 (North American genera key)
: 30 (Argentina genera); Menozzi & Russo, 1930Menozzi, C.; Russo, G. 1930. Contributo alla conoscenza della mirmecofauna della Repubblica Dominicana (Antille). Bollettino del Laboratorio di Zoologia Generale e Agraria della Reale Scuola Superiore d'Agricoltura. Portici 24:148-173. PDF
: 169 (Dominican Republic); Smith, 1937Smith, M. R. 1937 ("1936"). The ants of Puerto Rico. Journal of Agriculture of the University of Puerto Rico 20:819-875. [1937-01] PDF
: 822 (Puerto Rico); Kusnezov, 1956aKusnezov, N. 1956a. Claves para la identificación de las hormigas de la fauna argentina. Idia 104-105:1-56. [1956-09] PDF
: 7 (Argentina); Kusnezov, 1960bKusnezov, N. 1960b ("1959"). La fauna de hormigas en el oeste de la Patagonia y Tierra del Fuego. Acta Zoologica Lilloana 17:321-401. [1960-06-10] PDF
: 330 (West Patagonia); Snelling & Hunt, 1975Snelling, R. R.; Hunt, J. H. 1975. The ants of Chile (Hymenoptera: Formicidae). Revista Chilena de Entomología 9:63-129. [1976-05] PDF
: 63 (Chile); Kempf, 1978aKempf, W. W. 1978a. Formicidae-Hym. 111. Chave para identificação de operárias des sete subfamilias que ocorrem no Brasil. Studia Entomologica 20:33. [1978-08-30]
: 33 (Neotropical subfamilies); Baroni Urbani, 1984Baroni Urbani, C. 1984 ("1983"). Clave para la determinación de los géneros de hormigas neotropicales. Graellsia 39:73-82. [1984-02-10] PDF
: 73 (Neotropical genera); Mackay & Mackay, 1989: 1 (Mexico genera); Cover, in Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 69 (Neotropical genera key); Lattke, in Jaffe, 1993Jaffe, K. 1993. El mundo de las hormigas. Baruta, Venezuela: Equinoccio (Ediciones de la Universidad Simón Bolívar), 188 pp. [1993-07]
: 145 (Neotropical subfamilies & genera); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 26, 38, 49, 98, 162 (Dolichoderinae, Ecitoninae, Formicinae, Myrmicinae, Ponerinae genera keys); Fernández, Palacio, MacKay & MacKay, 1996Fernández, F.; Palacio, E. E.; MacKay, W. P.; MacKay, E. S. 1996. Introducción al estudio de las hormigas (Hymenoptera: Formicidae) de Colombia. Pp. 349-412 in: Andrade, M. G.; Amat García, G.; Fernández, F. (eds.) 1996. Insectos de Colombia. Estudios escogidos. Bogotá: Academia Colombiana de Ciencias Exactas, Físicas y Naturales, 541 pp. [1996-09]
: 353 (Neotropical subfamilies); Fernández & Ospina, in Fernández, 2003Fernández, F. 2003. Contributions in Introdución a las hormigas de la región Neotropical. 299-306, 307-330, 371-378, 379-411 in: Fernández, F. (ed.) 2003. Introducción a las hormigas de la región Neotropical. Bogotá: Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, xxvi + 424 pp.
: 49 (Neotropical genera synopsis); Palacio & Fernández, in Fernández, 2003Fernández, F. 2003. Contributions in Introdución a las hormigas de la región Neotropical. 299-306, 307-330, 371-378, 379-411 in: Fernández, F. (ed.) 2003. Introducción a las hormigas de la región Neotropical. Bogotá: Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, xxvi + 424 pp.
: 233 (Neotropical subfamilies and genera keys)
: 8 (Madagascar genera); Arnold, 1915Arnold, G. 1915. A monograph of the Formicidae of South Africa. Part I. Ponerinae, Dorylinae. Annals of the South African Museum 14:1-159. [1915-02-11]
: 6 (South Africa); Arnold, 1916Arnold, G. 1916. A monograph of the Formicidae of South Africa. Part II. Ponerinae, Dorylinae. Annals of the South African Museum 14:159-270. [1916-05-20] PDF
: 159 (South Africa); Bolton, 1973aBolton, B. 1973a. The ant genera of West Africa: a synonymic synopsis with keys (Hymenoptera: Formicidae). Bulletin of the British Museum (Natural History). Entomology 27:317-368. [1973-01-18] PDF
: 321 (West Africa subfamilies & genera); Bolton, in Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 39 (Afrotropical and Malagasy genera key); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 23, 44, 79, 155 (Dolichoderinae, Formicinae, Myrmicinae, Ponerinae genera keys); Yoshimura & Fisher, 2007Yoshimura, M.; Fisher, B. L. 2007. A revision of male ants of the Malagasy region (Hymenoptera: Formicidae): key to subfamilies and treatment of the genera of Ponerinae. Zootaxa 1654:21-40. [2007-12-05] PDF
: 27 (Malagasy subfamilies, males key)
: 219 (India & Sri Lanka); Bingham, 1903Bingham, C. T. 1903. The fauna of British India, including Ceylon and Burma. Hymenoptera, Vol. II. Ants and Cuckoo-wasps. London: Taylor and Francis, 506 pp. [1903] PDF
: 1 (India, Sri Lanka & Burma); Wilson & Taylor, 1967bWilson, E. O.; Taylor, R. W. 1967b. The ants of Polynesia (Hymenoptera: Formicidae). Pacific Insects Monograph 14:1-109. [1967-08-15] PDF
: 10 (Polynesia); Bolton, in Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 46 (Oriental and Indo-Australian genera key); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 23, 45, 84, 158 (Dolichoderinae, Formicinae, Myrmicinae, Ponerinae genera keys); Wu & Wang, 1995aWu, J.; Wang, C. 1995a. The ants of China. [In Chinese.]. Beijing: China Forestry Publishing House, x + 214 pp. [1995-06]
: 30 (China); Zhou, 2001AZhou, S. 2001A. Ants of Guangxi. [In Chinese.]. Guilin, China: Guangxi Normal University Press, 255 pp. [2001-09-01] PDF
: 21 (China, Guangxi subfamilies); Clouse, 2007BClouse, R. M. 2007B. The ants of Micronesia (Hymenoptera: Formicidae). Micronesica 39:171-295. [2007-07]
: 190 (Micronesia); Framenau & Thomas, 2008Framenau, V. W.; Thomas, M. L. 2008. Ants (Hymenoptera: Formicidae) of Christmas Island (Indian Ocean): identification and distribution. Records of the Western Australian Museum 25:45-85. [2008] PDF
: 52 (Christmas I.); Terayama, 2009 (Taiwan); Eguchi, Bui & Yamane, 2011Eguchi, K.; Bui, T. V.; Yamane, S. 2011. Generic synopsis of the Formicidae of Vietnam (Insecta: Hymenoptera), part I — Myrmicinae and Pseudomyrmecinae. Zootaxa 2878:1-61. [2011-05-13] PDF
: 7 (Vietnam subfamilies key); Pfeiffer, et al. 2011: 32 (ants of Borneo, checklist); General & Alpert, 2012General, D. M.; Alpert, G. D. 2012. A synoptic review of the ant genera (Hymenoptera, Formicidae) of the Philippines. ZooKeys 200:1-111. [2012-06-05] PDF
: 68 (Philippines genera key); Marcus, 1953Marcus, H. 1953. Estudios mirmecológicos. Folia Universitaria. Cochabamba 6:17-68. [1953]
: 1 (Fiji)
: 353 (New Zealand); Brown, 1958h: 1 (New Zealand); Brown & Taylor, 1970: 958 (Australia subfamilies); Greenslade, 1979Greenslade, P. J. M. 1979. A guide to ants of South Australia. Adelaide: South Australian Museum (Special Educational Bulletin Series), 44 pp. [1979]
: 10 (South Australia subfamilies); Taylor, R.W., in Hölldobler & Wilson, 1990Hölldobler, B.; Wilson, E. O. 1990. The ants. Cambridge, Mass.: Harvard University Press, xii + 732 pp.
: 55 (Australia genera key); Taylor, 1991dTaylor, R. W. 1991d. Formicidae. Pp. 980-989 in: CSIRO (ed.) 1991. The insects of Australia. Second edition. Volume II. . Carlton, Victoria: Melbourne University Press, pp. i-vi, 543-1137. [1991]
: 987 (Australia subfamilies); Bolton, 1994Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. [1994-07-25]
: 24, 47, 91, 159 (Dolichoderinae, Formicinae, Myrmicinae, Ponerinae genera keys); Shattuck, 1999Shattuck, S. O. 1999. Australian ants. Their biology and identification. Collingwood, Victoria: CSIRO Publishing, xi + 226 pp. [1999-01]
: 21 (Australia subfamilies & genera); Andersen, 2000Andersen, A. N. 2000. The ants of northern Australia. A guide to the monsoonal fauna. Collingwood, Victoria: CSIRO Publishing, v + 106 pp.
: 14 (northern Australia subfamilies & genera); Don, 2007Don, W. 2007. Ants of New Zealand. Dunedin, New Zealand: Otago University Press, 239 pp. [2007]
: 39 (New Zealand); Heterick, 2009Heterick, B. E. 2009. A guide to the ants of south-western Australia. Records of the Western Australian Museum Supplement 76:1-206. PDF
: 16, 28 (south-western Australia subfamilies & genera)| Genera |
| Species |
| †fossil | valid | synonym | subspecies | unidentifiable/excluded from Formicidae | unavailable/nomen nudum | homonym | original combination | View full key |